Cloth $35.00 ISBN: 9780226728346 Will Publish November 2020
E-book $35.00 Available for pre-order. ISBN: 9780226728483 Will Publish November 2020

Yellowstone Wolves

Science and Discovery in the World’s First National Park

Edited by Douglas W. Smith, Daniel R. Stahler, and Daniel R. MacNulty

Yellowstone Wolves

Edited by Douglas W. Smith, Daniel R. Stahler, and Daniel R. MacNulty

With a Foreword by Jane Goodall
344 pages | 62 color plates, 29 halftones, 24 line drawings | 8-1/2 x 11
Cloth $35.00 ISBN: 9780226728346 Will Publish November 2020
E-book $35.00 ISBN: 9780226728483 Will Publish November 2020
In 2020, it will have been twenty-five years since one of the greatest wildlife conservation and restoration achievements of the twentieth century took place: the reintroduction of wolves to the world’s first national park, Yellowstone. Eradicated after the park was established, then absent for seventy years, these iconic carnivores returned to Yellowstone in 1995 when the US government reversed its century-old policy of extermination and—despite some political and cultural opposition—began the reintroduction of forty-one wild wolves from Canada and northwest Montana. In the intervening decades, scientists have studied their myriad behaviors, from predation to mating to wolf pup play, building a one-of-a-kind field study that has both allowed us to witness how the arrival of top predators can change an entire ecosystem and provided a critical window into impacts on prey, pack composition, and much else.
Here, for the first time in a single book, is the incredible story of the wolves’ return to Yellowstone National Park as told by the very people responsible for their reintroduction, study, and management. Anchored in what we have learned from Yellowstone, highlighting the unique blend of research techniques that have given us this knowledge, and addressing the major issues that wolves still face today, this book is as wide-ranging and awe-inspiring as the Yellowstone restoration effort itself. We learn about individual wolves, population dynamics, wolf-prey relationships, genetics, disease, management and policy, newly studied behaviors and interactions with other species, and the rippling ecosystem effects wolves have had on Yellowstone’s wild and rare landscape. Perhaps most importantly of all, the book also offers solutions to ongoing controversies and debates.
Featuring a foreword by Jane Goodall, beautiful images, a companion online documentary by celebrated filmmaker Bob Landis, and contributions from more than seventy wolf and wildlife conservation luminaries from Yellowstone and around the world, Yellowstone Wolves is a gripping, accessible celebration of the extraordinary Yellowstone Wolf Project—and of the park through which these majestic and important creatures once again roam.
Study Area Map

A Note on Accompanying Video
Robert K. Landis

Jane Goodall

Douglas W. Smith, Daniel R. Stahler, and Daniel R. MacNulty

Part 1 History and Reintroduction
1 Historical and Ecological Context for Wolf Recovery
Douglas W. Smith, Daniel R. Stahler, Daniel R. MacNulty, and Lee H. Whittlesey

Box 1.1 Wolf History and Surveys in Yellowstone National Park
John Weaver

2 How Wolves Returned to Yellowstone
Steven H. Fritts, Rebecca J. Watters, Edward E. Bangs, Douglas W. Smith, and Michael K. Phillips

Box 2.1 To Reintroduce or Not to Reintroduce, That Is the Question
Diane Boyd

Guest Essay: Why Are Yellowstone Wolves Important?
L. David Mech

Part 2 Behavioral and Population Ecology

3 Essential Biology of the Wolf: Foundations and Advances
Daniel R. MacNulty, Daniel R. Stahler, Tim Coulson, and Douglas W. Smith

4 Ecology of Family Dynamics in Yellowstone Wolf Packs
Daniel R. Stahler, Douglas W. Smith, Kira A. Cassidy, Erin E. Stahler, Matthew C. Metz, Rick McIntyre, and Daniel R. MacNulty

Box 4.1 Naming Wolf Packs
Daniel R. Stahler

5 Territoriality and Competition between Wolf Packs
Kira A. Cassidy, Douglas W. Smith, Daniel R. Stahler, Daniel R. MacNulty, Erin E. Stahler, and Matthew C. Metz
Box 5.1 Auditory Profile: The Howl of the Wolf 
John B. Theberge and Mary T. Theberge

6 Population Dynamics and Demography
Douglas W. Smith, Kira A. Cassidy, Daniel R. Stahler, Daniel R. MacNulty, Quinn Harrison, Ben Balmford, Erin E. Stahler, Ellen E. Brandell, and Tim Coulson

Guest Essay: Yellowstone Wolves Are Important Because They Changed Science
Rolf O. Peterson and Trevor S. Peterson

Part 3 Genetics and Disease

7 Yellowstone Wolves at the Frontiers of Genetic Research
Daniel R. Stahler, Bridgett M. vonHoldt, Elizabeth Heppenheimer, and Robert K. Wayne

8 The K Locus: Rise of the Black Wolf
Rena M. Schweizer, Daniel R. Stahler, Daniel R. MacNulty, Tim Coulson, Phil Hedrick, Rachel Johnston, Kira A. Cassidy, Bridgett M. vonHoldt, and Robert K. Wayne

9 Infectious Diseases in Yellowstone’s Wolves
Ellen E. Brandell, Emily S. Almberg, Paul C. Cross, Andrew P. Dobson, Douglas W. Smith, and Peter J. Hudson

Guest Essay: Why Are Yellowstone Wolves Important? A European Perspective
Olof Liberg

Part 4 Wolf-Prey Relationships

10 How We Study Wolf-Prey Relationships
Douglas W. Smith, Matthew C. Metz, Daniel R. Stahler, and Daniel R. MacNulty
Box 10.1 Nine-Three-Alpha
Douglas W. Smith
Box 10.2 The Bone Collectors
Ky Koitzsch and Lisa Koitzsch

11 Limits to Wolf Predatory Performance 
Daniel R. MacNulty, Daniel R. Stahler, and Douglas W. Smith
Box 11.1 Tougher Times for Yellowstone Wolves Reflected in Tooth Wear and Fracture
Blaire Van Valkenburgh

12 What Wolves Eat and Why
Matthew C. Metz, Mark Hebblewhite, Douglas W. Smith, Daniel R. Stahler, Daniel R. MacNulty, Aimee Tallian, and John A. Vucetich

Box 12.1 Bison in Wood Buffalo National Park
L. N. Carbyn

13 Wolf Predation on Elk in a Multi-Prey Environment
Matthew C. Metz, Douglas W. Smith, Daniel R. Stahler, Daniel R. MacNulty, and Mark Hebblewhite
Box 13.1 Generalizing Wolf-Prey Dynamics across Systems: Yellowstone, Banff, and Isle Royale
Mark Hebblewhite
Box 13.2 The Predator’s Perspective: Biomass of Prey
Matthew C. Metz

Box 13.3 Lessons from Denali National Park: Stability in Predator-Prey Dynamics Is a Pause on the Way to Somewhere Else
Layne Adams

14 Population Dynamics of Northern Yellowstone Elk after Wolf Reintroduction
Daniel R. MacNulty, Daniel R. Stahler, Travis Wyman, Joel Ruprecht, Lacy M. Smith, Michel T. Kohl, and Douglas W. Smith
Box 14.1 Wolves and Elk in the Madison Headwaters
Robert A. Garrott, P. J. White, Claire Gower, Matthew S. Becker, Shana Drimal, Ken L. Hamlin, and Fred G. R. Watson
Box 14.2 Ecology of Fear
Daniel R. Stahler and Daniel R. MacNulty

Guest Essay: The Value of Yellowstone’s Wolves? The Power of Choice
Michael K. Phillips

Part 5 Ecosystem Effects and Species Interactions

15 Indirect Effects of Carnivore Restoration on Vegetation
Rolf O. Peterson, Robert L. Beschta, David J. Cooper, N. Thompson Hobbs, Danielle Bilyeu Johnston, Eric J. Larsen, Kristin N. Marshall, Luke E. Painter, William J. Ripple, Joshua R. Rose, Douglas W. Smith, and Evan C. Wolf
Box 15.1 Long-Term Trends in Beaver, Moose, and Willow Status in the Southern Portion of the Absaroka-Beartooth Wilderness
Daniel B. Tyers

16 Competition and Coexistence among Yellowstone’s Meat Eaters
Daniel R. Stahler, Christopher C. Wilmers, Aimee Tallian, Colby B. Anton, Matthew C. Metz, Toni K. Ruth, Douglas W. Smith, Kerry A. Gunther, and Daniel R. MacNulty

Guest Essay: Old Dogs Taught Old Lessons
Paul C. Paquet

Part 6 Conservation, Management, and the Human Experience

17 Wolves and Humans in Yellowstone
Douglas W. Smith, Daniel R. Stahler, Rick McIntyre, Erin E. Stahler, and Kira A. Cassidy

18 The Wolf Watchers
Nathan Varley, Rick McIntyre, and James Halfpenny
Box 18.1 Bob Landis’s Yellowstone Wolves Documentaries 000

Box 18.2 Seeing Wolves
Robert Hayes

19 Conservation and Management: A Way Forward
Douglas W. Smith, P. J. White, Daniel R. Stahler, Rebecca J. Watters, Kira A. Cassidy, Adrian Wydeven, Jim Hammill, and David E. Hallac

Guest Essay: Making Better Sense of Wolves
Susan G. Clark
Rebecca J. Watters, Douglas W. Smith, Daniel R. Stahler, and Daniel R. MacNulty


Appendix: Species Names Used in the Text

Literature Cited

List of Contributors

Author Index

Subject Index
Review Quotes
Kirkus Reviews
"A gathering of scientific essays on the natural history of the reintroduced wolf population in Yellowstone National Park. Wolves were long a key predator in the Greater Yellowstone Ecosystem before they were extirpated. In 1995, they were reintroduced in what the editors characterize as 'a change of heart.' That change of heart is 'the single most important fact of wolf recovery everywhere.' . . . A welcome contribution to the conservation-biology literature on wolves."
Jane Goodall, from the foreword
Yellowstone Wolves summarizes over two decades of hard work, involving dozens of dedicated scientists and advocates, to bring these wolves back to Yellowstone. . . . Their voices are skillfully combined to tell the many-faceted narratives in this marvelous book. . . . The overall success of this long-term effort provides information that will be of inestimable value to other restoration projects, sharing methods that can help wolves and humans coexist in a changing world and an example of what can happen if people unite to give Mother Nature a chance.”
Mark W. Moffett, author of "The Human Swarm: How Our Societies Arise, Thrive, and Fall"
“Smith and his colleagues have graced us with the most comprehensive and elegant book ever written about the Yellowstone wolves. They take us deep into the human dimensions of wolf recovery while also giving us exceptional insights about social behavior not only for wolves, but for animal societies in general.”
Bernd Heinrich, author of "Mind of the Raven," "The Homing Instinct," and "White Feathers"
“I can’t praise this book enough. From the foreword by Jane Goodall to David Mech at the end, to all the now nearly lifetime work of researchers dedicated to this grand ecological experiment, important conservation effort, and inspirational project, Yellowstone Wolves is a model of success that will not only endure, but grow and sprout hope and inspiration globally. Bravo!”
James A. Estes, author of "Serendipity: An Ecologist’s Quest to Understand Nature"
“Wolves are surrounded by conflict and controversy, the substance of which has been dimmed by a veil of scientific uncertainty. Yellowstone Wolves is a readable and authoritative account about this iconic species and magical place, written by the scientists, managers, and conservationists who did the work. Read this book and learn what these on-the-ground professionals really know and think about wolves in the Yellowstone ecosystem.”
James M. Peek, professor emeritus, Department of Fish and Wildlife Sciences, University of Idaho
“I consider the restoration of the gray wolf into the northwestern United States to be the single most important event in the past fifty years of wildlife conservation and management. This volume will serve as a major reference for years as it covers the key social/ecological/behavioral/management issues related to the wolf in Yellowstone National Park. Virtually all of the principal individuals who have been involved in wolf investigations and management across the continent have contributed.”
Dean Cluff, Department of Environment and Natural Resources, Government of the Northwest Territories, Canada
“Very informative. Every reader of this book will learn something new and interesting about wolves, their ecology, or how they compare to other species in the animal world. It will be a great resource, sought out as the de facto authority on wolf knowledge and discovery for Yellowstone most certainly, but also on how the knowledge gained lends itself to understanding wolves elsewhere. Compelling.”
Mark Boyce, Department of Biological Sciences, University of Alberta, Canada
Yellowstone Wolves is a Herculean attempt to synthesize research on Yellowstone National Park subsequent to wolf recovery. It is likely to be touted as the definitive work on the subject for years. This is a necessary, wonderful book.”
New York Times
"Reintroducing the wolf to Yellowstone is arguably the world’s greatest wildlife experiment. The wolves’ progress has been documented meticulously by a team of hiking, driving and flying biologists and passionate volunteers—so much so that no wolf study comes close to yielding its abundance of information. The research generated has been distilled into a new book, Yellowstone Wolves, assembled by three of the biologists who studied the wolves’ return."
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